Eutheria (//; from Greek εὐ-, eu- "good" or "right" and θηρίον, thēríon "beast" hence "true beasts") is one of two mammalian clades with extant members that diverged in the Early Cretaceous or perhaps the Late Jurassic. All placental mammals are eutherians. Extant eutherians, their last common ancestor, and all extinct descendants of that ancestor are members of Placentalia.
|Juramaia, the oldest known eutherian|
Eutherians are distinguished from noneutherians by various phenotypic traits of the feet, ankles, jaws and teeth. All extant eutherians lack epipubic bones, which are present in all other living mammals (marsupials and monotremes). This allows for expansion of the abdomen during pregnancy.
Distinguishing features are:
- an enlarged malleolus ("little hammer") at the bottom of the tibia, the larger of the two shin bones
- the joint between the first metatarsal bone and the entocuneiform bone (the outermost of the three cuneiform bones) in the foot is offset farther back than the joint between the second metatarsal and middle cuneiform bones—in metatherians these joints are level with each other
- various features of jaws and teeth
Eutheria contains several extinct genera as well as larger groups, many with complicated taxonomic histories still not fully understood. Members of the Adapisoriculidae, Cimolesta and Leptictida have been previously placed within the out-dated placental group Insectivora, while Zhelestids have been considered primitive ungulates. However, more recent studies have suggested these enigmatic taxa represent stem group eutherians, more basal to Placentalia.
|The fossil eutherian species believed to be the oldest known is Juramaia sinensis, which lived about . Montanalestes was found in North America, while all other nonplacental eutherian fossils have been found in Asia. The earliest-known placental fossils have also been found in Asia.|
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|Wikispecies has information related to Eutheria|
|Wikisource has the text of the 1911 Encyclopædia Britannica article Monodelphia.|