List of informally named dinosaurs

  (Redirected from Bayosaurus)

This list of informally named dinosaurs is a listing of dinosaurs (excluding Aves; birds and their extinct relatives) that have never been given formally published scientific names. This list only includes names that were not properly published ("unavailable names") and have not since been published under a valid name. The following types of names are present on this list:

  • Nomen nudum, Latin for "naked name": A name that has appeared in print but has not yet been formally published by the standards of the International Commission on Zoological Nomenclature. Nomina nuda (the plural form) are invalid, and are therefore not italicized as a proper generic name would be.
  • Nomen manuscriptum, Latin for "manuscript name": A name that appears in manuscript but was not formally published. A nomen manuscriptum is equivalent to a nomen nudum for everything except the method of publication, and description.
  • Nicknames or descriptive names given to specimens or taxa by researchers or the press.
"Nurosaurus" mounted skeleton, Inner Mongolia Museum



"Airakoraptor" is an informal name given to a genus of dromaeosaurian theropod from the Upper Cretaceous of Mongolia. The nomen nudum was accidentally created by Perle et al. in their 1992 in a citation for a paper called "Morphology Dromaeosaurian dinosaur-Airakoraptor from the upper cretaceous of Mongolia".[1][2] As the journal and volume of the listed paper contain no such publication, the actual title for the work likely being "New dromaeosaur material from the Late Cretaceous of Mongolia" by the same authors as listed, there is no verifiable publication for this taxon name. The proper publication described three specimens, one of which is not referred to by a specimen number, and most likely corresponds to the material "Airakoraptor" was to be named for. This specimen, IGM 100/981, was found in Khulsan and is referrable to Dromaeosauridae while likely distinct from Velociraptor.[2]


"Amargastegos" is an informally named genus of extinct stegosaurid ornithischian dinosaur known from the La Amarga Formation of Argentina on the basis of MACN N-43 (some dorsal osteoderms, the cervical and caudal vertebrae, and one skull bone). In 2016, Peter Malcolm Galton and Kenneth Carpenter declared it a nomen dubium, establishing it as an indeterminate stegosaur.[3]


"Andhrasaurus indicus" is an informal genus of extinct armored ornithischian dinosaur from the Kota Formation of India. Ulansky (2014) coined the name for skull elements, about 30 osteoderms, and the extremities of vertebrae and limbs, all preserved in the collections of the GSI and assigned to Ankylosauria by Nath et al. (2002).[4] In 2016, Peter Malcolm Galton and Kenneth Carpenter noted that "Andhrasaurus" did not meet ICZN requirements and therefore declared it a nomen nudum, listing it as Thyreophora indet., while noting that the jawbones described by Nath et al. (2002) belonging to crocodylomorphs.[3] The dermal armor informally named "Andhrasaurus" was redescribed by Galton (2019).[5]

Angeac ornithomimosaurEdit

The "Angeac ornthomimosaur" is an informal name given to an unnamed orthithomimosaur taxon known from an Early Cretaceous (previously thought to be Hauterivian-Barremian in age,[6] but now thought to be Berriasian-Valanginian based on palynology[7]) bone bed (part of the stratigraphy of the Aquitaine Basin) near Angeac in France. The taxon is toothless and is known from multiple individuals collectively covering much of the skeleton, some remains were described by Allain et al. (2014).[8]


Pneumatic structures of "Angloposeidon"

"Angloposeidon" is the informal name given to a sauropod dinosaur from the Early Cretaceous Wessex Formation of the Isle of Wight in southern England.[9] It was a possible brachiosaurid but has not been formally named. Darren Naish, a notable vertebrate palaeontologist, has worked with the specimen and has recommended that this name only be used informally and that it not be published.[10] However, he published it himself in his book Tetrapod Zoology Book One from 2010.[11] The remains consist of a single cervical vertebra (MIWG.7306), which indicate it was a very large animal, 20 metres or greater in length.[12]


The "Archaeoraptor" fossil

"Archaeoraptor" is the informal generic name for an important fossil from China that was later discovered to be a fake. The name was created in an article published in National Geographic magazine in 1999, where the magazine claimed that the fossil was a "missing link" between birds and terrestrial theropod dinosaurs. Even prior to this publication there had been severe doubts about the fossil's authenticity. Further scientific study showed it to be a forgery constructed from rearranged pieces of real fossils from different species. Zhou et al. found that the head and upper body actually belong to a specimen of the primitive fossil bird Yanornis, and another 2002 study found that the tail belongs to a small winged dromaeosaur, Microraptor, named in 2000.[13][14] The legs and feet belong to an as yet unknown animal.[15][16]


"The Archbishop" in multiple views

"The Archbishop" is a giant brachiosaurid sauropod dinosaur similar to Brachiosaurus and Giraffatitan. It was long considered a specimen of Brachiosaurus (now Giraffatitan) brancai due to being found in the same formation in Tendaguru, Tanzania. However, the "Archbishop" shows significant differences including a unique vertebral morphology and a proportionally longer neck, that indicates it is a different, previously unknown genus and species.[17] It was discovered by Frederick Migeod in 1930. "The Archbishop" is a nickname that functions as a placeholder – the specimen currently has no scientific name. The specimen is currently housed in the Natural History Museum in London, and will eventually be re-described by Dr. Michael P. Taylor of Bristol University.[18] In May 2018, Taylor started to work on describing the Archbishop.[19]



"Balochisaurus" (meaning "Balochi lizard", for the Baloch tribes of Pakistan) is an informal taxon of titanosaurian sauropod dinosaur from the Late Cretaceous of Pakistan. The proposed species is "B. malkani". The discovery was made (along with other dinosaur specimens) near Vitariki by a team of paleontologists from the Geological Survey of Pakistan.[20] Described in 2006 by M.S. Malkani, the genus is based on seven tail vertebrae found in the Maastrichtian-age Vitakri Member of the Pab Formation, with additional vertebrae and a partial skull assigned to it. Balochisaurus was assigned to the family "Balochisauridae" along with "Marisaurus", although the family was used as a synonym of older Saltasauridae.[21]


"Bananabendersaurus"[22] is the informal name given to an unnamed species of sauropod found in the Winton Formation, Australia.[23] The two known specimens of "Bananabendersaurus", known as Cooper and George, were discovered in 2005 and 2006 and have been dated to the early Cenomanian (ca. 98-95 Ma).[24] The Cooper specimen is known from both humeri and the left radius, while the George specimen is known from a single rib and a left femur.[24] "Bananabendersaurus" represents the largest dinosaur discovered in Australia to date - "Bananabendersaurus" was around 26 metres (85 ft) long on average.


"Barackosaurus" is the informal name created in 2010 which is used for a sauropod found in Kimmeridgian-aged sediments pertaining to the Morrison Formation, Wyoming. It was found in the Dana Quarry and "Barackosaurus" was supposedly 20 meters long and weighed 20 tons.[23]

Barnes High SauropodEdit

The "Barnes High sauropod" is the informal name given to MIWG-BP001, an undescribed sauropod dinosaur specimen from the Wessex Formation on the Isle of Wight. It was discovered in the cliffs around Barnes High in 1992 and is currently owned by the privately run unaccredited Dinosaur Farm Museum near Brighstone,[25] the ownership situation was described as "complex" and the specimen is currently inaccessible to researchers.[26] It is roughly 40% complete and consists of a "Partial postcranial skeleton, including presacral vertebrae, anterior caudal vertebrae, girdle and limb elements" including a largely complete forelimb. It has been suggested to be a Brachiosaur and is possibly synonymous with the earlier named Eucamerotus due to similarities with the vertebrae.[27]


"Bayosaurus" is the informal name given to an as yet undescribed genus of dinosaur. The name was coined by paleontologists Rodolfo Coria, Philip J. Currie, and Paulina Carabajal in 2006. It apparently was an abelisauroid from the Turonian Cerro Lisandro Formation of Neuquén, Argentina, around 4 m (13 ft) long. The specimen is MCF-PVPH-237, including dorsal and sacral vertebrae, a fragmentary pelvis, and other partial bones, which were discovered in 2000. The name was used in a phylogenetic analysis to indicate the position of MCF-PVPH-237.[28] The suggested binomial was "Bayosaurus pubica".


"Beelemodon" is the informal name given to an undescribed theropod genus from the Late Jurassic, possibly belonging to a coelurosaur. The fossils include two teeth found in Wyoming, United States. The name appeared in print in 1997, when paleontologist Robert T. Bakker mentioned it in a symposium for the Academy of Natural Sciences.[29][30] The teeth are most similar to Compsognathus, but have no unique features and also share similarities with Protarchaeopteryx and dromaeosaurids.[30]


"Biconcavoposeidon" is the placeholder name for AMNH FARB 291, five consecutive posterior dorsal vertebrae of a brachiosaurid sauropod, from the Late Jurassic Morrison Formation, Wyoming.[31] Not much else is currently known about "Biconcaveoposeidon", except that it was discovered in the Bone Cabin quarry in 1898.[32]


"Bihariosaurus" (meaning "Bihor lizard") is an invalid genus of iguanodontian dinosaur from Early Cretaceous Bauxite of Cornet, Romania. The type species, "Bihariosaurus bauxiticus", was named but not described by Marinescu in 1989. It was similar to Camptosaurus, and was an iguanodont. The original publication of the taxon did not include sufficient description, and the illustrations cannot distinguish it from any other ornithopod.[33][34] "Bihariosaurus" may have been the same animal as Valdosaurus.


"Biscoveosaurus" is the informal name of an ornithopod dinosaur specimen from the Early Maastrichtian age Snow Hill Island Formation of James Ross Island, Antarctica. It comes from the Cape Lamb Member of the formation, the same member as Morrosaurus, another basal ornithopod. As such, it's been suggested it may be a secondary specimen of that species, but as the holotype of Morrosaurus is fragmentary and doesn't overlap with the material of "Biscoveosaurus", this can't as yet be tested. The specimen consists of dentaries, teeth, a braincase, parts of the maxillae, forelimb elements, assorted vertebrae, and the pectoral girdle; this makes it unique compared to the other James Ross Island ornithopods, which don't presever both cranial and postcranial remains. It has been estimated the animal would've been about 4–5 metres (13–16 ft) in length.[35]

Brachiosaurus nougarediEdit

Diagram showing preserved parts of the "B." nougaredi sacrum in blue

"Brachiosaurus" nougaredi was a sauropod dinosaur of uncertain affinities. It was originally assigned to the genus Brachiosaurus in 1960, though it certainly represents a different genus, and probably a different family.

In 1958, the French petroleum geologist F. Nougarède reported to have discovered fragmentary brachiosaurid remains in eastern Algeria, in the Sahara Desert.[36] Based on these, Albert-Félix de Lapparent described and named the species Brachiosaurus nougaredi in 1960. He indicated the discovery locality as being in the Late Jurassic–age Taouratine Series. He assigned the rocks to this age in part because of the presumed presence of Brachiosaurus.[37] A more recent review placed it in the "Continental intercalaire," which is considered to belong to the Albian age of the late Early Cretaceous, significantly younger.[38]

The type material moved to Paris consisted of a sacrum, weathered out at the desert surface, and some of the left metacarpals and phalanges. Found at the discovery site but not collected, were partial bones of the left forearm, wrist bones, a right shin bone, and fragments that may have come from metatarsals.[37]

"B." nougaredi was in 2004 considered to represent a distinct, unnamed brachiosaurid genus,[38] but a 2013 analysis by Philip D. Mannion and colleagues found that the remains possibly belong to more than one species, as they were collected far apart.[39] The metacarpals were concluded to belong to some indeterminate titanosauriform. The sacrum was reported lost in 2013. It was not analyzed and provisionally considered to represent an indeterminate sauropod, until such time that it could be relocated in the collections of the Muséum national d'histoire naturelle. Only four out of the five sacral vertebrae are preserved. The total original length was in 1960 estimated at 1.3 meters (4 ft 3 in), compared to 0.91 meters (3 ft 0 in) with B. altithorax.[37] This would make it larger than any other sauropod sacrum ever found, except those of Argentinosaurus and Apatosaurus.[39]



"Capitalsaurus" vertebra

"Capitalsaurus" is the informal genus name given to the species formerly known as Creosaurus potens and Dryptosaurus potens, a dinosaur from the Arundel Formation of North America. The only known specimen was discovered in Washington, D.C. It was a theropod, and it lived during the Cretaceous. It is a nomen nudum, the name having never been formally published. "Capitalsaurus" was discovered in January 1898 at the intersection of First and F Streets S.E., in the District of Columbia – an intersection now designated as Capitalsaurus Court. It was not uncovered by any paleontological activity, but as a by-product of sewer work. The only known specimen of "Capitalsaurus" consists of part of a single vertebra. Some paleontologists feel that this is insufficient justification for a name that suggests an entire genus, and that "Capitalsaurus" is merely an undetermined theropod.[40] Others note that this is hardly the only dinosaur with a common name that does not helpfully reflect its taxonomy.[41] The "Capitalsaurus" is the official dinosaur of the District of Columbia.[42]


"Changdusaurus" (also known as "Changtusaurus") is the informal name given to a genus of dinosaur from the Late Jurassic Period. It lived in what is now China. "Changdusaurus" is classified as a stegosaurid. The type species was named "Changdusaurus laminoplacodus" by Zhao in 1986,[43] but it has never been formally described, and remains a nomen nudum. One source indicates the fossils have been lost.[44]


"Coelurosaurus" is an informal generic name, attributed to Friedrich von Huene, 1929, that is sometimes seen in lists of dinosaurs. It probably arose as a typographical error; von Huene intended to assign indeterminate remains to Coelurosauria incertae sedis, but at some point in the process of publication a revision to the text made it appear that he was creating a new generic name "Coelurosaurus" (as described by George Olshevsky in a 1999 post to the Dinosaur Mailing List). The name is undescribed and has not been used seriously, although it has appeared in works of fiction.


"Comanchesaurus" is an informal name for fossilized remains from the Late Triassic of New Mexico that were initially interpreted as belonging to a theropod dinosaur. The remains, NMMNH P-4569, consist of a partial skeleton including vertebral centra and hindlimb bones, and came from the Norian-age Upper Triassic Bull Canyon Formation of Guadalupe County. Adrian Hunt, in his unpublished dissertation, proposed the name "Comanchesaurus kuesi" for the specimen, but the name was never adopted, and was first referred to in the scientific literature in a 2007 redescription of Late Triassic North American material thought to belong to dinosaurs (Nesbitt, Irmis, and Parker, 2007). In the redescription, the authors found the material to belong to a "possible indeterminate saurischian".[45]



"Dachongosaurus" is the informal name given to an undescribed genus of sauropod dinosaur from the Early Jurassic of China. It is known from fossils including at least a partial articulated skeleton from the Dark Red Beds of the Lower Lufeng Series (Sinemurian stage) in Yunnan.[46] Possibly a cetiosaur, the "type species" is "Dachongosaurus yunnanensis", coined by Zhao in 1985. An alternate spelling is "Dachungosaurus". As with other informal names coined by Zhao in 1985 and 1983, nothing has since been published, and the remains may have been redescribed under another name.[47]


"Damalasaurus" (meaning "Damala lizard") is the informal name given to a genus of herbivorous dinosaur from the Early Jurassic. It was a sauropod, though its exact classification within the clade is unknown. Fossils of "Damalasaurus", including a rib, have been found in the Middle Daye Group of Tibet. Species attributed to this genus include "Damalasaurus laticostalis" and "D. magnus".[46]


"Duranteceratops" is a purported new taxon of chasmosaurine ceratopsid from the Hell Creek Formation.[48] In 2012, a ceratopsid skull reportedly distinguishable from Triceratops was unearthed in South Dakota by an amateur fossil hunter named John Carter.[49][50][48] Though it has yet to be published, according to the Prehistoric Times issue no. 121 from Spring 2017, the specimen is to be named "Duranteceratops".


EK troodontidEdit

The "EK troodontid" (specimen SPS 100/44) is an unnamed genus of troodontid dinosaur discovered in Mongolia. In the scientific literature it is referred to as the "EK troodontid", after the Early Cretaceous sediments in which it was found. SPS 100/44 was discovered by S.M. Kurzanov during the 1979 Soviet-Mongolian Paleontological Expedition. It was found in deposits of the Barunbayaskaya Svita at the Khamareen Us locality, Dornogov (southeastern Gobi Desert), in the Mongolian People's Republic. SPS 100/44 was described by Rinchen Barsbold and colleagues in 1987.[51]

Its fossil remains include an incomplete skeleton consisting of the braincase, posterior parts of the lower mandibles, a maxillary fragment with teeth, parts of five cervical vertebrae, an articulated right manus with partial semilunate, left manus phalanx I-1, distal end of the left femur, and fragmentary left and right pedes. Barsbold pointed out that the specimen was smaller and from older sediments than other known troodontids, but it had some features of the skull that could have made it a juvenile. Barsbold also indicated the high degree of fusion of the bones of the skull and the unusual foot morphology to indicate that it might be an adult of an unknown taxon. Barsbold took the conservative position and did not name this specimen because it was not complete enough to rule out the possibility that it was a juvenile of a known genus of troodontid. Barsbold also noted that the naturally articulated manus of SPS 100/44 showed no signs of an opposable third digit, as was suggested for Troodon by Russell and Seguin in 1982. Turner and colleagues, in 2007, found the EK troodontid to be a distinct basal genus of troodontid, in a polytomy with Jinfengopteryx and a clade of more derived troodontids.[52]


"Eoplophysis" is a genus of stegosaur known from the Middle Jurassic Cornbrash Formation, Sharp's Hill Formation, and Chipping Norton Formation of England.[53] It was originally named Omosaurus vetustus by the renowned German paleontologist Friedrich von Huene.[54] The holotype, OUM J.14000, is a 60-centimetre-long (2 ft) right femur of a juvenile individual from the Middle Jurassic (upper Bathonian) Cornbrash Formation of Oxfordshire, England, although it was probably reworked from the slightly older Forest Marble Formation in view of its eroded nature. Because of the renaming of Omosaurus, an occupied name, as Dacentrurus, O. vetustus was renamed into a Dacentrurus vetustus in 1964.[55] In the 1980s, researcher Peter Malcolm Galton reviewed all known stegosaur material from the Bathonian of England and concluded that Omosaurus vetustus was valid and should be tentatively referred to Lexovisaurus.[56][57] However, the species was later considered a nomen dubium in both reviews of Stegosauria.[58][59] In their alpha-taxonomic review of stegosaurs, Susannah Maidment and her colleagues noted that OUM J.14000 shares characters present in both sauropods and stegosaurs, but that it lacks synapomorphies exclusive to Stegosauria and assigned it as a Dinosauria indet.[60] Nevertheless, the amateur paleontologist Roman Ulansky coined the new genus "Eoplophysis" ("Dawn Armed Form") for O. vetustus, noting differences with the femora of other stegosaurs.[61]


"Eugongbusaurus" skull IVPP 14559

"Eugongbusaurus" is the informal name (nomen nudum) proposed for a neornithischian found in the Oxfordian-age Shishugou Formation of Xinjiang, China. The intended type species, "Gongbusaurus" wucaiwanensis, was described by Dong Zhiming in 1989 for two partial skeletons as a second species of the poorly known tooth taxon Gongbusaurus. Fragmentary skeleton IVPP 8302, the type specimen for the new species, included a partial lower jaw, three tail vertebrae, and a partial forelimb. Second specimen IVPP 8303 consisted of two hip vertebrae, eight tail vertebrae, and two complete hind limbs. Dong estimated it as around 1.3 to 1.5 meters (4.3 to 4.9 ft) long, and considered it to be a strong runner. He assigned the genus Gongbusaurus to the Hypsilophodontidae, a paraphyletic grade of small herbivorous bipedal dinosaurs.[62] Because dinosaur teeth are generally not distinctive enough to hold a name, it is unsurprising that other paleontologists have suggested removing "G." wucaiwanensis from Gongbusaurus and giving it its own genus.[63] The possible replacement name "Eugongbusaurus"[64] leaked out accidentally and remains informal.



Fendusaurus is a nomen ex dissertatione proposed by Fedak (2006) for FGM998GF13-II, which includes a skull. Other specimens referred to Fendusaurus are FGM998GF13-I, FGM998GF13-III, FGM998GF69, FGM998GF9, and FGM998GF18, all found by a crew from the Princeton University. All the specimens include femora and coracoids, and although they each share slightly different features, the differences are credited to intra-specific variation. Known specimens of Fendusaurus were previously classified as cf. Ammosaurus. The femora and coracoids also help identify different individuals, and Timothy J. Fedak, the described of the specimens, found that each block represented about one individual. Fendusaurus is known from the Early Jurassic (Hettangian) McCoy Brook Formation of Wasson Bluff, Nova Scotia. As five specimens of Fendusaurus are from the McCoy Brook Formation, the formation is the richest prosauropod site in North America. The formation is also similar to other formations of North America and Asia, as it lacks any remains presently assigned to Anchisaurus. Fedak places Fendusaurus as a genus of the family Massospondylidae.[65]

The specimens of Fendusaurus include mostly crushed vertebrae, along with appendicular elements. They are distinguishable from Anchisaurus by the morphology of both the ilium and sacral vertebrae. However, in some specimens, the morphology of the femora and coracoids are quite different, which led Fedak to speculate that more than one species may have been present. Fendusaurus, according to Fedak, can be distinguished from all closely related sauropodomorphs by the extreme elongation of the cervical vertebrae; a four vertebrae sacrum that includes a dorsosacral and caudosacral; the elongate postacetabular process of the ilium; and an expanded anterior distal process of the tibia.[65]


"Ferganastegos" is a dubious genus of stegosaur from the Middle Jurassic (Callovian) Balabansai Formation of Fergana Valley, Kyrgyzstan.[66][67] The holotype of "Ferganastegos callovicus", IGB 001, consists of four posterior dorsal vertebrae.[68] Although Averianov et al. did not consider the vertebrae diagnostic to genus, the freelance Russian dinosaur enthusiast and amateur paleontologist Roman Ulansky decided that the differences between IGB 001 and other stegosaurs were sufficient to warrant a binomial for IGB 001, "Ferganastegos callovicus" (Callovian roof from Fergana Valley), despite the fact he did not examine the material himself.[66] Other researchers still contend that the material is not diagnostic and that the genus is a nomen dubium.[69]


"Futabasaurus" is an informal name for a genus of theropod dinosaur from the Late Cretaceous of Japan, known only from a partial shin bone discovered in the Coniacian-age Ashizawa Formation of the Futaba Group.[70] It was coined by David Lambert in 1990 as a conversion from the Japanese nickname "Futaba-ryu", for an undescribed theropod.[71] Dong Zhiming and coauthors briefly discussed the fossil shin bone it was based on that same year, publishing a photograph. They considered the bone to belong to an indeterminate tyrannosaurid.[72] If the specimen is eventually described and named, it will require a different name, because the name Futabasaurus has since been used for a genus of plesiosaur.[73]



Skeleton of "Gadolosaurus"

"Gadolosaurus" is an informal name given to an undescribed hadrosauroid from the Bayan Shireh Formation of Baishan Tsav, Mongolia.[74] The name "Gadolosaurus" was first used in a 1979 book by Japanese paleontologist Tsunemasa Saito, in a caption for a photo of a juvenile dinosaur skeleton.[75] This small individual was only about a meter long (39 inches). The skeleton was part of a Soviet exhibition of fossils in Japan. Apparently, the name comes from a Japanese phonetic translation of the Cyrillic word gadrosavr, or hadrosaur, and was never meant by the Russians to establish a new generic name.[76]

Despite being merely a mistranslation of gadrosavr, it appeared in many popular dinosaur books, with varying identifications. Donald F. Glut in 1982 reported it as either an iguanodont or hadrosaur, with no crest or boot on the ischium (the lack of which are both characteristics of the crested lambeosaurine duckbills), and suggested it could be the juvenile of a previously named genus like Tanius or Shantungosaurus.[77] David Lambert in 1983 classified it as an iguanodont,[78] but changed his mind by 1990, when it was listed as a synonym of Arstanosaurus without comment.[71] What may be the same animal is mentioned but not named by David B. Norman and Hans-Dieter Sues in a 2000 book on Mesozoic reptiles from Mongolia and the former USSR; this material, from the Soviet-Mongolian expeditions of the 1970s, had been listed as Arstanosaurus in the Russian Academy of Sciences, and was found in the Cenomanian-age Bayan Shireh Formation of Baishin Tsav.[79]

Gallimimus mongoliensisEdit

"Gallimimus mongoliensis" skeleton

"Gallimimus mongoliensis" is an informal name Rinchen Barsbold used for a nearly complete skeleton (IGM 100/14) known from the Bayan Shireh Formation, but since it differs from Gallimimus in some details, Yoshitsugu Kobayashi and Barsbold proposed in 2006 that it probably belongs to a different genus.[80]


"Gspsaurus" (a nomen manuscriptum) is a titanosaurian sauropod dinosaur from the Late Cretaceous Vitakri Formation of Sulaiman Basin of Pakistan.[81] It has been suggested to be synonymous with the also invalid taxon "Maojandino", also proposed by Malkani.[82][83]


"Grusimimus" (or "Tsurumimus") is an informal name for an undescribed genus of ornithomimid from the Early Cretaceous (HauterivianBarremian) aged Shinekhudag Formation of Mongolia. Known from a skeleton including all regions except the skull, "Grusimimus" was given an invalid name in 1997 by Rinchen Barsbold, who also suggested the species name "tsuru". The specimen (GIN 960910KD) was found in 1996 and examined by Barsbold before he suggested the informal name, a nomen nudum. An abstract and poster were presented on the taxon by Kobayashi & Barsbold in 2002, and the former published a thesis paper on the specimen (referred to as "Ornithomimosauria indet.") which found the taxon to be close to Harpymimus phylogenetically but possible more derived.[84]



"Hanwulosaurus" is the informal name given to an as-yet undescribed genus of dinosaur from the Late Cretaceous. It was an ankylosaur around 9 m (30 ft) long, which is long for an ankylosaur. Its fossils were found in Inner Mongolia, China. Much of a skeleton, including a complete skull, vertebrae, ribs, a scapula, an ulna, femorae, bones from the shin, and armor, was discovered; this may be the most complete ankylosaurian skeleton yet found in Asia, according to early reports. Zhao Xijin, who has studied it, suggests that it may belong to its own subgroup within the Ankylosauria. The name first surfaced in news reports in 2001.[citation needed]


"Heilongjiangosaurus" (meaning "Heilongjiang lizard") is the informal name given to an as-yet undescribed genus of duckbilled dinosaur from the Late Cretaceous. It possibly was a lambeosaurine, and may in fact be the same animal as Charonosaurus. The fossils were found in Maastrichtian-age rocks in Heilongjiang, China. As a nomen nudum, it is unclear what material it was intended to be based on, but might be connected to the nomen nudum "Mandschurosaurus" jiainensis,[85] informally named in a 1983 publication.[86]

The "type species" is "H. jiayinensis", and it was coined in 2001 in a faunal list by Li and Jin.[87]


"Hironosaurus" (meaning "Hirono lizard") is the informal name given to an as-yet undescribed genus of dinosaur from the Late Cretaceous. Found in Hirono, Fukushima, Japan, it was probably a type of hadrosaur, although no subfamily identification has been made. The fossils are quite fragmentary, and consist of teeth and a vertebra, possibly from the tail. Since the fossils have never been fully described in a scientific paper, "Hironosaurus" is considered a nomen nudum. It was first mentioned by Hisa in an obscure 1988 publication[88] and was later (1990) brought to a wider audience by David Lambert.[71] Dong Zhiming, Y. Hasegawa, and Y. Azuma regarded the material as belonging to a hadrosaurid, but lacking any characteristics to allow more precise identification (thus indeterminate).[72]


"Hisanohamasaurus" (meaning "Hisano-hama lizard") is the informal name given to an as yet undescribed genus of dinosaur from the Late Cretaceous. It is a nomen nudum known only from teeth that first appeared in a general-audience dinosaur book by David Lambert in 1990. Although initially identified a diplodocid,[89] it later re-identified as a nemegtosaurid similar to Nemegtosaurus.[90] As its name suggests, its fossils were found in Japan. The location is part of Iwaki, Fukushima.



"Ichabodcraniosaurus" is an informal name for theropod fossils were first discovered within the Gobi Desert in China. The remains consist of a partial skeleton without a head (IGM 100/980), from which its name refers to the fictional character Ichabod Crane, a character in the history of Washington Irving's The Legend of Sleepy Hollow, where he was pursued by a headless horseman. Fossils include a hyoid bone, cervical vertebrae, a supposed skull and a supposed jaw bone. "Ichabodcraniosaurus" was named by Novacek in 1996.[91] This specimen may belong to Velociraptor mongoliensis, but Norell and Makovicky concluded that it was not complete enough to say for sure, and it awaits a formal description.[92]



"Jiangjunmiaosaurus" is an informal name created by an anonymous author in 1987 for a possible chimaera of Monolophosaurus and Sinraptor.[93]


"Jindipelta" (Lei et al., 2019; in press) is the currently informal name given to an ankylosaur from the Zhumapu Formation in China. It is known from a partial skeleton found in Cenomanian rocks. The name was first announced in the 2019 SVP abstract book, alongside the megalosauroid Yunyangosaurus.[94]


Cast of Julieraptor

"Julieraptor" is the nickname of a dromaeosaurid fossil found in the Judith River Formation, Montana in 2002. Parts of the same skeleton were illegally excavated and nickamed Sid Vicious in 2006, and the excavator subsequently served jail time for the theft. Bob Bakker therefore also nicknamed the specimen "Kleptoraptor". The skeleton was arranged to be sold to Royal Ontario Museum.[95][96][97] It is known from a skeleton consisting of an almost complete skeleton missing most of its skull, most tail vertebra, part of the femur, some spinal and neck vertebra, one claw but it has a well preserved braincase.



"Kagasaurus" (meaning "Kaga lizard") is the informal name given to an as yet undescribed genus of dinosaur from the Early Cretaceous. It was a theropod which lived in what is now Japan. The type species was named by Hisa in 1988, but is known from only two teeth. Since "Kagasaurus" has never been formally described, it is considered a nomen nudum. Unlike "Kitadanisaurus" and Katsuyamasaurus, it is unlikely that "Kagasaurus" is synonymous with Fukuiraptor, and may instead be a dromaeosaurid.[citation needed]


"Katsuyamasaurus" is an informal name for a genus of intermediate theropod known from the Early Cretaceous (Barremian) of the Kitadani Formation, Japan. Known from a single middle caudal vertebra and an ulna, the taxon was informally called "Katsuyama-ryu", until Lambert (1990) made it into an invalid genus name, "Katsuyamasaurus". The caudal vertebra was suggested to belong to an ornithopod by Chure (2000), and Olshevsky (2000) suggested the material was a synonym of Fukuiraptor. However, the ulna differs from Fukuiraptor, and the large olecranon suggests the taxon falls outside Maniraptoriformes.[98]

Kelmayisaurus gigantusEdit

"Kelmayisaurus gigantus" is an informal name for a genus of sauropod known from the Cretaceous. The type species of Kelmayisaurus, K. petrolicus, was first named and described by Chinese paleontologist Dong Zhiming in 1973.[99] A supposed second species, K. "gigantus", was mentioned by Grady in a children book in 1993 as a gigantic vertebral column coming from a 22 m long specimen,[100] but is a nomen nudum and probably does not pertain to Kelmayisaurus. It may be a sauropod instead.[101]


"Khetranisaurus" (meaning "Khetran lizard", for the Khetran people of Pakistan) is an informal taxon of titanosaurian sauropod from the Late Cretaceous of Balochistan, western Pakistan (also spelled "Khateranisaurus" in some early reports).[102] The proposed species is "K. barkhani", described by M. Sadiq Malkani in 2006, and it is based on a tail vertebra, found in the Maastrichtian-age Vitakri Member of the Pab Formation. It was assigned to "Pakisauridae" (used as a synonym of Titanosauridae), along with "Pakisaurus" and "Sulaimanisaurus".[21]


"Koreanosaurus" (meaning "Korean lizard") is the informal name given to an as-yet undescribed genus of dinosaur from the Early Cretaceous. It was a possible dromaeosaur (or similar theropod) which lived in what is now Korea, although at times it has been referred to the Tyrannosauridae and Hypsilophodontidae. Based solely on a femur, the name was coined by Kim in 1979, but by 1993 Kim decided that it was a species of Deinonychus, and created the informal name "D." "koreanensis".[103]


"Kunmingosaurus" is an informally named primitive sauropod which lived during the Early Jurassic. Its fossils were found in Yunnan Province, China in 1954. The type and only species is "Kunmingosaurus wudingensis", invalidly coined by Zhao in 1985. It is known from fossils found in the Fengjiahe Formation (or the Lower Lufeng Series), including pelvic, hind limb, and vertebral material.[46][104][105][106]



"Lancanjiangosaurus" (alternative spelling "Lanchanjiangosaurus"; meaning "Lancangjiang lizard", named after the Lancangjiang River of China) is the informal name given to an as yet undescribed genus of sauropod dinosaur from the Middle Jurassic. The "type species", "L. cachuensis", was coined by Zhou in 1985, but remains a nomen nudum. It is known from the Dapuka Group of Tibet.[46]


"Lijiagousaurus" (meaning "Lijiagou lizard") is the informal name given to an as yet undescribed genus of herbivorous iguanodontian dinosaur from the Late Cretaceous of what is now Sichuan, China. It has not been formally described yet, but the formal publication is forthcoming, from Chinese paleontologist Ouyang Hui. "Lijiagousaurus" was only briefly mentioned in the Chongqing Natural History Museum guidebook (2001) and is thus a nomen nudum.[107][108]


"Likhoelesaurus" (meaning "Li Khole lizard") is the name given to an as yet undescribed genus of archosauriform, either a dinosaur or rauisuchian, from the Late Triassic of what is now South Africa. The name was coined by Ellenberger in 1970, and the "type species" is "Likhoelesaurus ingens".[109] It is named after the town in Lesotho where the fossils were found. The only fossils recovered have been teeth, from the late Carnianearly Norian-age Lower Elliot Formation.[110] Ellenberger (1972) regarded the genus as a giant carnosaur, and Kitching and Raath (1984) treated it as possibly referable to Basutodon.[111][112] Knoll listed "Likhoelesaurus" as a rauisuchian, also he noted that could also be a rauisuchian.[113]



"Magulodon" is the name given to an as yet undescribed genus of dinosaur from the Early Cretaceous (Aptian to Albian stages, approximately 112 million years ago). It was a possible ornithischian, either an ornithopod or basal ceratopsian, which lived in what is now Maryland, in the United States. The type species, "Magulodon muirkirkensis", was coined by Kranz in 1996.[114] It is a tooth taxon, based solely on a single tooth. Since it has not been formally described, it is also a nomen nudum. It was considered to be an indeterminate specimen in a paper which cited the intended type specimen but avoided using the name to prevent taxonomic clutter.[115]


"Marisaurus" (meaning "Mari lizard", for the Mari tribe of Pakistan) is an informal taxon of titanosaurian sauropod from the Late Cretaceous of Balochistan, western Pakistan. The type species is "M. jeffi", described by M. Sadiq Malkani in 2006, and it is based on tail vertebrae, found in the Maastrichtian-age Vitakri Member of the Pab Formation. Much additional material, including a partial skull, many vertebrae, and a few hindlimb bones, was referred to this genus. "Marisaurus" was assigned to "Balochisauridae" with "Sulaimanisaurus", although the family was used as a synonym of Saltasauridae.[21][116]


"Megacervixosaurus" (meaning "big neck lizard") is the informal name given to an as yet undescribed genus of herbivorous dinosaur from the Late Cretaceous. It was a titanosaur sauropod which lived in what is now China. The type species, "Megacervixosaurus tibetensis", was coined by Chinese paleontologist Zhao Xijin in 1985. "Megacervixosaurus" has never been formally described, and remains a nomen nudum.[117]


Knee-joint and a claw of Merosaurus

"Merosaurus" is the informal name given to an indeterminate genus of dinosaur from the Early Jurassic (Sinemurian stage, around 190 million years ago) of Charmouth, Dorset It originates either from the Blue Lias Formation or the Charmouth Mudstone Formation. It was theropod, possibly a tetanuran which lived in what is now England. The type species, "Merosaurus newmani", was coined by paleontologists Samuel Welles, H.P. Powell, and A. Pickering in 1995, and is based solely on some leg bones (a knee joint) once thought to belong to Scelidosaurus.[118][119]


"Microcephale" (meaning "tiny head") is the informal name of a genus of very small pachycephalosaurid dinosaur, otherwise known as the "North American dwarf species", which lived during the Late Cretaceous. Its fossils were found in the late Campanian-age Dinosaur Park Formation, in Alberta, Canada. Not much is known about this dinosaur, as it has not yet been fully described; it is therefore a nomen nudum. The fossils of "Microcephale", including tiny skull caps, were first mentioned by paleontologist Paul Sereno in 1997, in a list of pachycephalosaurids. These skull caps measure less than 5 cm (2 in) each. No potential species name was given.


"Microdontosaurus" (meaning "tiny-toothed lizard") is the name given to an as yet undescribed genus of sauropod dinosaur from China. It was named from fossils from the Middle Jurassic-age Dapuka Group of Xinjiang. The intended type species is "M. dayensis."[46] As with other informal names created by Zhao in 1985 or 1983, it has not been used since then, and may have been redescribed under another name.[120]


"Mifunesaurus" (meaning 'Mifune lizard') is a nomen nudum given to an extinct non-avian theropod dinosaur from Cenomanian rocks of Japan. "Mifunesaurus" is only known from a few bones, among which are a tibia, a phalanx, a metatarsus and a single tooth. The genus was informally coined by Hisa in 1985.[121]


The "Moshisaurus" humerus

Hisa (1985) used "Moshisaurus" (or "Moshi-ryu") for sauropod remains from the Early Cretaceous Miyako Group of Japan. Dong et al. (1990) and Hasegawa et al. (1991) referred them to Mamenchisaurus, but Azuma & Tomida (1998) and Barrett et al. (2002) assigned them to Sauropoda indet.[122][123][124]



Illustration of the "Newtonsaurus" dentary mold in internal and external views

"Newtonsaurus" is an informally named genus erected for the theropod dinosaur species Zanclodon cambrensis, which was discovered in the Lilstock Formation in 1898. It was probably a ceratosaur, which lived in what is now the United Kingdom. The taxon was reassigned to Megalosaurus due to the taxonomic difficulties associated with Zanclodon. It is based on a mold of a dentary from Rhaetian-age beds in Wales (hence the species name), and is one of the relatively few dinosaurs known from the time near the TriassicJurassic boundary.[125]

Paleontologists have avoided using the name "Newtonsaurus" since its appearance in 1999 in private publications, although "Zanclodon" cambrensis or Megalosaurus cambrensis have both been used for this taxon.[126][127]


"Ngexisaurus" is the informal name given to an as yet undescribed genus of theropod dinosaur from the Middle Jurassic Dapuka Group of Tibet, China. The type species, "Ngexisaurus dapukaensis", was coined by Zhao in 1983.[128][129][130]


"Nicksaurus" is an informally named Titanosaurian sauropod dinosaur from the Late Cretaceous red muds of the Vitakri Formation of Sulaiman Basin, Pakistan. The dinosaur shared a habitat with other sauropod dinosaurs including Khetranisaurus, Sulaimanisaurus, Pakisaurus, Gspsaurus, Saraikimasoom, and Maojandino.[131]


A close-up of the head of "Nurosaurus qaganensis".

"Nurosaurus" (Nur-o-saw-rus, meaning "Nur lizard") is the informal name for a genus of sauropod dinosaur. It is known from a partial, large skeleton, that was presented as soon-to-be-described by Zhiming Dong in 1992, where he gave the proposed binomial "Nurosaurus qaganensis". It was discovered in the Qagannur Formation of Inner Mongolia, 65 km (40 mi) southeast of Erenhot. The deposit is younger than the Psittacosaurus-bearing Guyang Group, but is still Early Cretaceous. It was found alongside the plates and scapula of a stegosaur.[132]

The foot of "Nurosaurus" is notable for a stress fracture present on the first phalanx of the fourth digit of the left foot, which was the first identified fracture of its kind, and have since been identified on the phalanges and metatarsals of Apatosaurus, Barosaurus, Brachiosaurus, Camarasaurus, and Diplodocus.[133]



"Orcomimus" (Pronounced or-coh-MEEM-us) is the name given to an as yet undescribed genus of dinosaur from the Late Cretaceous period 66 million years ago. The dinosaur was an ornithomimid which lived in what is now South Dakota, in the United States. The type was coined by Michael Triebold in 1997, but has never been formally described and is currently a nomen nudum. "Orcomimus" was a bipedal theropod, but the dinosaur is known from only a pelvis and a hindlimb. "Orcomimus" is thought to be relatively advanced for other ornithomimids at the time, although this is hard to tell from the limited amount of specimens found of the dinosaur.[134] It may be referable to one of the ornithomimosaur species currently known from the Hell Creek Formation.


"Oshanosaurus" (meaning "Oshan lizard") is the informal name given to an as yet undescribed genus of sauropod dinosaur from the Early Jurassic period of Yunnan, China. Its fossils were found in the Lower Lufeng Series. The intended "type species", "Oshanosaurus youngi", was coined by Zhao in 1985.[46] It has sometimes been associated with heterodontosaurids, which appears to be due to the juxtaposition of a species of Dianchungosaurus (formerly thought to be a heterodontosaurid) in the text of Zhao (1985).[135]



"Pakisaurus" (meaning "Pakistan lizard") is an informal taxon of titanosaurian sauropod from the Late Cretaceous of Balochistan, western Pakistan. The proposed species is "P. balochistani", and it was validly named by M. Sadiq Malkani in 2006, based on four tail vertebrae, found in the Maastrichtian-age Vitakri Member of the Pab Formation. Three additional tail vertebrae have been assigned to it. The author erected the family "Pakisauridae", using it synonymously with older Titanosauridae. "Pakisaurus" was decided to be closely related to "Sulaimanisaurus" and "Khetranisaurus", both from Pakistan as well.[21]


"Podischion" is an informal genus of hadrosaurid dinosaur known from a skeleton discovered in 1911 on the Red Deer River in Alberta by a crew led by Barnum Brown. The remains were tentatively named "Podischion", which was not mentioned in published literature until Dingus & Norell (2010).[136][93] It is possible that the skeleton represents an individual of Hypacrosaurus.[93]



Cast of "Ronaldoraptor"

"Ronaldoraptor", also known as the “Mitrata” Oviraptorid, is an undescribed oviraptorid from Mongolia[137] and has been listed as "Oviraptor sp."[138] The name was first used by Luis Rey in 2003, in his book A Field Guide to Dinosaurs: The Essential Handbook for Travelers in the Mesozoic, where he drew an illustration, captioning it "Ronaldoraptor".[137] "Ronaldoraptor" may have been closely related to Citipati osmolskae.


"Rutellum" is the pre-Linnaean name given to a dinosaur specimen from the Middle Jurassic. It was a sauropod, possibly a cetiosaurid,[139] which lived in what is now England. The specimen, called "Rutellum impicatum", was described in 1699 by Edward Lhuyd,[140] and is notable as the earliest named entity that is recognizable as a dinosaur.[139] It was based on a tooth collected from Caswell, near Witney, Oxfordshire.[141]

Because "Rutellum impicatum" was named before 1758 (the official starting date for zoological nomenclature according to the ICZN), it is not considered a part of modern biological nomenclature.



"Sabinosaurus" mount

"Sabinosaurus" or "Sabinosaurio" is a name used for a partial skeleton from the Sabinas Basin in Mexico. It was initially described as Kritosaurus sp. by Jim Kirkland and colleagues,[142] but considered an indeterminate saurolophine by Prieto-Márquez (2014).[143] This skeleton is about 20% larger than other known specimens, around 11 meters [36 ft] long, and with a distinctively curved ischium, and represents the largest known well-documented North American saurolophine. Unfortunately, the nasal bones are also incomplete in the skull remains from this material.[142]


"Saldamosaurus" is an informal genus of stegosaurid dinosaur known from a complete braincase discovered in the Early Cretaceous Saldam Formation of Siberia, Russia. The type species, "Saldamosaurus tuvensis", was named in 2014 but according to Galton and Carpenter (2016) it did not meet the requirements of the International Code of Zoological Nomenclature and is hence a nomen nudum,.[144][145]


Skeleton of "Saltillomimus"

"Saltillomimus" is an informal name for an ornithomimid theropod from the Late Cretaceous (late Campanian) of the Cerro del Pueblo Formation in Mexico. It is known from a partial tail, most of a hindlimb, and forelimb bones that was given the name "Saltillomimus rapidus" by Martinez in 2010. Named in his thesis, the taxon name is an invalid nomen ex dissertatione.[84]


"Sanchusaurus" (meaning "Lizard from Sanchu") is an informal name for an ornithomimosaur dinosaur from the Early Cretaceous period of Asia. It is only known by a partial tail vertebra,[72] found in Nakasato, Japan.[146] Dong (1990) [72] considered it synonymous with Gallimimus but the large discrepancy in both age and location between the two species renders this opinion untenable. The genus has not been formally described and is considered a nomen nudum. It was first mentioned by Hisa in 1985.[147]


"Saraikimasoom" (meaning 'Innocent one') is an invalid species of titanosaur dinosaur from the Vitakri Formation in Pakistan. The type species, Saraikimasoom vitakri, was described by Sadiq Malkhani in 2015, in a paper describing multiple Pakistani dinosaurs, such as Gspsaurus, "Nicksaurus" and "Maojandino".[148] Saraikimasoom is currently recognised as a nomen manuscriptum.

Shake-N-Bake theropodEdit

The "Shake-N-Bake theropod" is an undescribed species of coelophysoid from the Kayenta Formation.[149][150]


"Siamodracon" is an extinct genus of invalid stegosaurid dinosaur known from a single dorsal vertebra found in Thailand's Phu Kradung Formation. According to Galton and Carpenter (2016) it did not meet the requirements of the International Code of Zoological Nomenclature.[151][144] "Siamodracon" was the first thyreophoran dinosaur discovered in South East Asia.


"Sidormimus" is an informal genus of noasaurid discovered in the Elrhaz Formation in Niger. It was discovered in 2000 by Chris Sidor and it was immediately described by Lyon on the Project Exploration website, with a photograph of the holotype.[152] During the same year, on the National Geographic website, the same photograph of the holotype was labelled "Dongosaurus". In 2005, Sidor himself confirmed that "Sidormimus" was the Elrhaz noasaurid. "Sidormimus" has been mentioned by Paul Sereno three times.[153][154][155] "Sidormimus" is known from a partial post cranial skeleton. Its neck and ribs were exposed when the holotype was discovered.


"Sinopeltosaurus" is a dubious genus of extinct scelidosaurid ornithischian dinosaur described by Roman Ulansky. The type and only species is "S. minimus" of the lower Jurassic Lufeng Formation of Yunnan China, based on an articulated set of ankle bones. In 2016, Peter Malcolm Galton and Kenneth Carpenter identified it as a nomen dubium, and listed it as Ornithischia indet., possible Thyreophora indet.[3][61][156] Ulansky variously referred to it as "Sinopeltosaurus minimus" or "Sinopelta minima"; Galton and Carpenter, as the first revisers under ICZN, made the former official.[3]


Skeletal diagram of "Skaladromeus"

"Skaladromeus" or the "Kaiparowits ornithopod" is an ornithopod from the Kaiparowits Formation named in a 2012 thesis by Clint Boyd.[157][158] The intended type species is "Skaladromeus goldenii".


The upper femur of "Suciasaurus"

A fossil theropod nicknamed "Suciasaurus rex" was discovered in 2012 at Sucia Island State Park in San Juan County of the U.S. State of Washington. It was the first dinosaur discovered in Washington state.[159][160] The finding was announced when Burke Museum paleontologists published a discovery paper in PLoS ONE.[161][162] Prompted by a petition from students at an elementary school at Parkland, near Tacoma, the Washington State Legislature introduced a bill in 2019 to make it the official state dinosaur.[159][160][163]


"Sugiyamasaurus" (meaning "Sugiyama lizard") is the informal name given to a few spatulate teeth belonging to a titanosauriform, possibly Fukuititan, which lived in Japan during the Early Cretaceous. The name was first printed by David Lambert in 1990 in the Dinosaur Data Book, and also appears in Lambert's Ultimate Dinosaur Book and in many on-line lists of dinosaurs. Since it has not been formally described, "Sugiyamasaurus" is a nomen nudum. Remains were found near Katsuyama City and were initially referred to Camarasauridae, but might belong to Fukutitan because they were unearthed in the same quarry as the Fukuititan material.[164][165][166]


"Sulaimanisaurus" (meaning "Sulaiman lizard", for the Sulaiman foldbelt) is an informal taxon of titanosaurian sauropod from the Late Cretaceous of Balochistan, western Pakistan (also spelled "Sulaimansaurus" in some early reports).[102] The proposed species is "S. gingerichi", described by M. Sadiq Malkani in 2006, and it is based on seven tail vertebra, found in the Maastrichtian-age Vitakri Member of the Pab Formation. Four additional tail vertebrae have been assigned to it. It was considered to be related to "Pakisaurus" and "Khetranisaurus" in the family "Pakisauridae" (used as a synonym of Titanosauridae).[21]



Syntype tyrannosauroid tibia AMNH 2550 given the name "Teihivenator"

"Teihivenator" ("strong hunter") is an improperly named taxon of tyrannosauroid coelurosaur from the Navesink Formation of New Jersey. It was suggested to contain the species, "T." macropus, originally classified as a species of Dryptosaurus (= "Laelaps", a name preoccupied by a mite). It was suggested as a separate genus in 2017 by Chan-gyu Yun.[167] The name "Teihivenator" is invalid because the publication naming it is online-only, which means that a registration with ZooBank is required to be present in the article when published. However, the ZooBank registry was only added in after initial publication, meaning that it fails the requirement to be a validly published taxon.[168]

In 2017, a preprint paper by Chase Brownstein concluded that the remains of L. macropus are a mixture of tyrannosauroid and ornithomimid elements with no distinguishing characteristics, rendering the species a chimera and a nomen dubium.[169] In 2018, Brownstein stated that a tibia of L. macropus catalogued as specimen AMNH FARB 2550 represents a tyrannosauroid that probably was distinct from Dryptosaurus, but not sufficiently to base a taxon on.[170]


"Tiantaisaurus", alternatively spelled "Tiantaiosaurus", is the name given to a specimen of therizinosaur from the Aptian age Laijia Formation of Zhejiang Province, China. According to correspondence through the Dinosaur Mailing List, the former name (from a 2012 study) was the one intended to be use for an official description. After being discovered in 2005, it was first mentioned named in an unpublished manuscript written in 2007. The given species was named "T. sifengensis". The specimen consists of an ischium, an astragalus, a tibia, a femur, an incomplete pubis and ilium, and a large number of vertebrae from across the body.[171][172]


"Tonouchisaurus" (meaning "Tonouchi lizard") is the informal name given to an as yet undescribed genus of dinosaur from the Early Cretaceous Period. It was a theropod which lived in what is now Mongolia. The suggested "type species", "Tonouchisaurus mongoliensis", was coined by Barsbold in 1994. It was notably small: less than 0.91 m (3 ft) in length. It may have been a tyrannosaurid, judging by the known skeletal elements, which include only limb bones. These limb bones include "a complete didactyl manus and complete pes, as well as other limb bones" (Olshevsky 1996).[173]


Valdosaurus dextrapodaEdit

In 1998 William Blows inadvertently named another species of Valdosaurus, "V. dextrapoda", by including this name in a fauna list,[174] but this was an error, and the species has never been supported.[175][176] Lacking description, it is a nomen nudum.


In 1982 Justin Delair informally named the genus "Vectensia" based on specimen GH 981.45, an armour plate. Like the holotype of Polacanthus it was found at Barnes High, but reportedly in an older layer, of the Lower Wessex Formation.[177] Blows in 1987 tentatively referred it to Polacanthus.[178]


"Vitakridrinda" is a genus of abelisaurid theropod dinosaur from the Late Cretaceous of Balochistan, western Pakistan. The intended type species is "V. sulaimani". The discovery was made (along with other dinosaur specimens) near Vitariki by a team of palaeontologists from the Geological Survey of Pakistan, in rocks from the Maastrichtian-age Vitakri Member of the Pab Formation.[21] Formally described in 2006 by M.S. Malkani, the genus is based on partial remains including two thigh bones, a braincase, and a tooth. A partial snout was later found that Malkani assigned to the holotype, and additional vertebrae may also belong to this genus. However, the snout was later reclassified as a new genus of mesoeucrocodylian, Induszalim.[179][180][181] Thomas Holtz gave a possible length of 6 meters (19.7 feet).[182] It is, however, a possible nomen nudum.[183]


"Vitakrisaurus" is a genus of noasaurid theropod dinosaurs represented by only one known species, "Vitakrisaurus saraiki", which is the intended type species. It lived in the late Cretaceous period, approximately 70 million years ago, during the Maastrichtian, in what is today the Indian subcontinent. Its fossils were found in Pakistan's Vitraki Formation. The holotype specimen, MSM-303-2 is a right foot with a seemingly tridactyl form and robust phalanges. It may belong to Noasauridae due to similarities with the foot of Velocisaurus, although inconsistencies within its brief description and a lack of comparison with other theropods within the article makes formal classification difficult.

The generic name references the Vitakri Member of the Pab Formation and combines this with the Greek suffix "saurus", meaning "reptile". The specific name honours the Saraiki people, who primarily live in southern Pakistan. However, like most dinosaur taxa named by M. Sadiq Malkani, it is probably a nomen nudum[183]



Skeletal mount of "Xinghesaurus" from a Japanese fossil expo

"Xinghesaurus" was the name given to a species of sauropod dinosaur, possibly a titanosauriform, in 2009, in the guidebook for the dinosaur expo "Miracle of Deserts". No species name was given for the genus.[184][185]



"Yuanmouraptor" or "Yuenmeuraptor" is an informally named carnosaur from Yuanmou County, China.[186] It lived during the Middle Jurassic, around 174 and 163 million years ago, and it is known from ZLJ0115, which is a complete, articulated skull on display at an unknown Chinese museum (possibly the Lufeng Dinosaur Museum), alongside a reconstructed skeleton of "Yuanmouraptor". "Yuanmouraptor" was briefly mentioned in a 2014 guide book.


"Yibinosaurus" (meaning "Yibin lizard") is the informal name given to an as yet undescribed genus of herbivorous dinosaur from the Early Jurassic. It was a sauropod which lived in what is now Sichuan, China. The suggested "type species", "Yibinosaurus zhoui", has not been formally described yet, but the formal publication is forthcoming, from Chinese paleontologist Ouyang Hui. "Yibinosaurus" was only briefly mentioned in the Chongqing Natural History Museum guidebook (2001) and is thus a nomen nudum.[107][108]


"Yunxianosaurus" is the provisional name for a genus of titanosaurian dinosaurs from the Late Cretaceous of what is now Hubei Province, China. The type species, "Yunxianosaurus hubeinensis", was proposed by Chinese paleontologist Li Zhengqi in 2001. The fossils of "Yunxianosaurus" were found near the Nanyang Prefecture. Li stated that the name "Yunxianosaurus" was a temporary label for ease of description, but that further field work and study of the fossils would be required before the genus could be given an official name.[187][188]


Zamyn Khondt oviraptoridEdit

Zamyn Khondt oviraptorid

Zamyn Khondt oviraptorid is a nickname for oviraptorid specimen IGM or GIN 100/42. Since the type skull of Oviraptor is so poorly preserved and crushed, the skull of IGM 100/42 has become the quintessential depiction of that dinosaur, even appearing in scientific papers with the label Oviraptor philoceratops.[189] However, this distinctive-looking, tall-crested species has more features of the skull in common with Citipati than it does with Oviraptor and it may represent a second species of Citipati or possibly an entirely new genus, pending further study.[190]

Zizhongosaurus huangshibanensisEdit

In 1999 Li Kui mentioned "Zizhongosaurus huangshibanensis"[191] but this has remained an undescribed nomen nudum.

See alsoEdit


  1. ^ Perle, A.; Norell, M.A.; Clark, J.M. (1999). "A new maniraptoran Theropod – Achillobator giganticus (Dromaeosauridae) – from the Upper Cretaceous of Burkhant, Mongolia". Contribution No. 101 of the Mongolian-American Paleontological Project: 1–105.
  2. ^ a b Mortimer, M. (2018). "Dromaeosaurs". TheropodDatabase. Retrieved 2018-07-16.
  3. ^ a b c d Galton, Peter M. & Carpenter, Kenneth, 2016, "The plated dinosaur Stegosaurus longispinus Gilmore, 1914 (Dinosauria: Ornithischia; Upper Jurassic, western USA), type species of Alcovasaurus n. gen.", Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen 279(2): 185-208
  4. ^ Nath, T. T., Yadagiri, P., and Moitra, A. K., 2002, First record of armoured dinosaur from the Lower Jurassic Kota Formation, Pranhita-Godavari Valley, Andhra Pradesh: Journal of the Geological Society of India, v. 59, p. 575-577.
  5. ^ Peter M. Galton (2019). "Earliest record of an ankylosaurian dinosaur (Ornithischia: Thyreophora): Dermal armor from Lower Kota Formation (Lower Jurassic) of India". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 291 (2): 205–219. doi:10.1127/njgpa/2019/0800.
  6. ^ Néraudeau, D.; Allain, R.; Ballèvre, M.; Batten, D.J.; Buffetaut, E.; Colin, J.P.; Dabard, M.P.; Daviero-Gomez, V.; El Albani, A.; Gomez, B.; Grosheny, D. (October 2012). "The Hauterivian–Barremian lignitic bone bed of Angeac (Charente, south-west France): stratigraphical, palaeobiological and palaeogeographical implications". Cretaceous Research. 37: 1–14. doi:10.1016/j.cretres.2012.01.006.
  7. ^ Polette, France; Batten, David J.; Néraudeau, Didier (October 2018). "Re-examination of the palynological content of the Lower Cretaceous deposits of Angeac, Charente, south-west France: Age, palaeoenvironment and taxonomic determinations" (PDF). Cretaceous Research. 90: 204–221. doi:10.1016/j.cretres.2018.04.017.
  8. ^ "European ornithomimosaurs (Dinosauria, Theropoda): an undetected record" (PDF). Geologica Acta (17). 2014. doi:10.1344/105.000002083.
  9. ^ Naish, Darren (2007-12-10). "World first: a peek inside "Angloposeidon"". Sauropod Vertebra Picture of the Week. Retrieved 2018-07-17.
  10. ^ Naish, Darren (15 Jul 2006). "Angloposeidon', the unreported story, part IV". Darren Naish: Tetrapod Zoology.
  11. ^ Naish, Darren (7 Oct 2010). "Tetrapod Zoology Book One is here at last". Archived from the original on 2012-05-08.
  12. ^ Naish, D.; Martill, D.M.; Cooper, D.; Stevens, K.A. (2004). "Europe's largest dinosaur? A giant brachiosaurid cervical vertebra from the Wessex Formation (Early Cretaceous) of southern England" (PDF). Cretaceous Research. 25 (6): 787–795. doi:10.1016/j.cretres.2004.07.002.
  13. ^ Zhou, Z.; Clarke, J.A.; Zhang, F. (2002). "Archaeoraptor's better half". Nature. 420 (6913): 285. Bibcode:2002Natur.420..285Z. doi:10.1038/420285a. PMID 12447431. S2CID 4423242.
  14. ^ Mayell, H. (2002-11-20). "Dino Hoax Was Mainly Made of Ancient Bird, Study Says". National Geographic. Archived from the original on 12 May 2008. Retrieved 2008-06-13.
  15. ^ Holden, C. (2000). "Florida Meeting Shows Perils, Promise of Dealing for Dinos". Science. 288 (5464): 238–239. doi:10.1126/science.288.5464.238a. PMID 10777394. S2CID 26985016.
  16. ^ Rowe, T.; Ketcham, R.A.; Denison, C.; Colbert, M.; Xing, X.; Currie, P.J. (2001). "Forensic palaeontology: The Archaeoraptor Forgery". Nature. 410 (6828): 539–540. Bibcode:2001Natur.410..539R. doi:10.1038/35069145. PMID 11279483. S2CID 205015615.
  17. ^ Specific features of the Archbishop that distinguish it from other brachiosaurs
  18. ^ "Just write the Archbishop description already!". 3 September 2015. Retrieved 1 January 2019.
  19. ^ "Work on the Archbishop begins!". 28 May 2018. Retrieved 1 January 2019.
  20. ^ Rana, A.N. (2006-03-25). "Country's first dinosaur fossils". DAWN. Archived from the original on 2007-06-21. Retrieved 2007-07-29.
  21. ^ a b c d e f Malkani, M.S. (2006). "Biodiversity of saurischian dinosaurs from the Latest Cretaceous Park of Pakistan" (PDF). Journal of Applied and Emerging Sciences. 1 (3): 108–140. Archived from the original (PDF) on 2009-06-22. Cite error: The named reference "MSM06" was defined multiple times with different content (see the help page).
  22. ^
  23. ^ a b
  24. ^ a b "Cooper & George". Queensland Museum. Retrieved 30 September 2020.
  25. ^ Naish, Darren (July 12, 2006). "'Angloposeidon', the unreported story, part I". Tetrapod Zoology.
  26. ^ Taylor, Mike (November 6, 2008). "Mystery sauropod dorsals of the Wealden, part 3: BMNH R88/89 — Britain's Best Brachiosaur (for now)". SVPOW.
  27. ^ "Sauropod dinosaurs". Field Guide to English Wealden Fossils. Palaeontological Association. 2011.
  28. ^ Coria, R.A; Currie, P.J.; Carabajal, A.P. (2006). "A new abelisauroid theropod from Northwestern Patagonia". Canadian Journal of Earth Sciences. 43 (9): 1283–1289. Bibcode:2006CaJES..43.1283C. doi:10.1139/e06-025.
  29. ^ Bakker, R. Raptor family values: Allosaur parents brought great carcasses into their lair to feed their young. In "Dinofest International", Proceedings of a Symposium, Academy of Natural Sciences, eds Wolberg, Sump and Rosenberg, 51 – 63 (1997).
  30. ^ a b Mortimer, M. (2001-01-31). "Details on "Beelemodon"". Dinosaur Mailing List Archives. Retrieved 2018-07-17.
  31. ^ "Biconcaveoposeidon". Sauropod Vertebra Picture of the Week. Retrieved 30 September 2020.
  32. ^ Taylor, Michael P; Wedel, Mathew J (2017). "A unique Morrison-Formation sauropod specimen with biconcave dorsal vertebrae". doi:10.7287/peerj.preprints.3144v1. Cite journal requires |journal= (help)
  33. ^ Posmoşanu, E. (2003). "Iguanodontian dinosaurs from the lower Cretaceous Bauxite site from Romania" (PDF). Acta Paleontologica Romaniae. 4: 431–439.
  34. ^ F. Marinescu. 1989. Lentila de bauxita 204 de la Brusturi-Cornet (Jud. Bihor), zacamint fosilifer cu dinozauri [Bauxite lens 204 from Brusturi-Cornet (Bihor Co.), fossil excavation with dinosaurs]. Ocrotirea Naturii si a Mediului Inconjurator 33(2):125-133
  35. ^ Lamanna, Matthew C.; Case, Judd A.; Roberts, Eric M.; Arbour, Victoria M.; Ely, Ricardo C.; Salisbury, Steven W.; Clarke, Julia A.; Malinzak, D. Edward; West, Abagael R.; O'connor, Patrick M. (2019). "Late Cretaceous non-avian dinosaurs from the James Ross Basin, Antarctica: description of new material, updated synthesis, biostratigraphy, and paleobiogeography". Advances in Polar Science. 30 (3): 228–250. doi:10.13679/j.advps.2019.0007.
  36. ^ Lapparent, A.F. de; Claracq, P.; Nougarède, F. (1958). "Nouvelles découvertes de Vertébrés dans les séries continentales au Nord d'Edjelch (Sahara central)". Comptes Rendus Hebdomadaires des Séances de l'Académie des Sciences de Paris. 247: 2399–2402.
  37. ^ a b c de Lapparent, A. F. (1960). Translated by Carrano, Matthew. "Les dinosauriens du "continental intercalaire" du Sahara central"" [The dinosaurs of the "continental intercalaire" of the central Sahara] (PDF). Mémoires de la Société Géologic de France, Nouvelle Séries 88A (in French). 39 (1–6): 1–57.
  38. ^ a b Upchurch, P.; Barrett, P.M.; Dodson, P. (2004). "Sauropoda". In Weishampel, D.B.; Dodson, P.; Osmolska, H. (eds.). The Dinosauria, Second Edition. Univ of California Press, Berkeley. pp. 259–322. ISBN 978-0-520-24209-8.
  39. ^ a b Mannion, P. D.; Upchurch, Paul; Barnes, Rosie N.; Mateus, Octávio (2013). "Osteology of the Late Jurassic Portuguese sauropod dinosaur Lusotitan atalaiensis (Macronaria) and the evolutionary history of basal titanosauriforms" (PDF). Zoological Journal of the Linnean Society. 168: 98–206. doi:10.1111/zoj.12029.
  40. ^ "Paleobiology - Smithsonian National Museum of Natural History". Retrieved 1 January 2019.
  41. ^ "Re: Details on Capitalsaurus: Multiple musings..." Retrieved 1 January 2019.
  42. ^ "LexisNexis® Legal Resources". Retrieved 1 January 2019.
  43. ^ Chao, S. (1983). "Phylogeny and Evolutionary Stages of Dinosauria". Acta Palaeontologica Polonica. 28: 295–306.
  44. ^ Maidment, Susannah C.R.; Guangbiao Wei (2006). "A review of the Late Jurassic stegosaurs (Dinosauria, Stegosauria) from the People's Republic of China". Geological Magazine. 143 (5): 621–634. Bibcode:2006GeoM..143..621M. doi:10.1017/S0016756806002500. Retrieved 2008-06-29.
  45. ^ Nesbitt, Sterling J.; Irmis, Randall B.; Parker, William G. (2007). "A critical re-evaluation of the Late Triassic dinosaur taxa of North America". Journal of Systematic Palaeontology. 5 (2): 209–243. doi:10.1017/S1477201907002040. S2CID 28782207.
  46. ^ a b c d e f Zhao, 1985. The reptilian fauna of the Jurassic in China. Pages 286–289, 347 in Wang, Cheng and Wang (eds.). The Jurassic System of China. Geological Publishing House, Beijing.
  47. ^ George Olshevsky (16 Nov 1999). "Re: What are these dinosaurs". Archives of the Dinosaur Mailing List at The Cleveland Museum of Natural History. Retrieved 2007-01-29.
  48. ^ a b O' Connell, Max. "Dinosaur skull found in Buffalo likely a new species". Rapid City Journal. Retrieved 27 August 2015.
  49. ^ Biles, Jan. "Rare dinosaur skull being prepared for exhibition". Topeka Topeka Capital-Journal. Retrieved 7 September 2015.
  50. ^ "Super-sized Ceratopsian Skull Might be New Species". Everything Dinosaur. Retrieved 29 August 2015.
  51. ^ Barsbold, Rhinchen, Osmolska, Halszka, Kurzanov, S.M. (1987). "On a new troodontid (Dinosauria. Theropoda) from the Early Cretaceous of Mongolia". Acta Palaeontologica Polonica 32(1-2): 121-132
  52. ^ Turner, Alan H.; Pol, Diego; Clarke, Julia A.; Erickson, Gregory M.; and Norell, Mark (2007) "A basal dromaeosaurid and size evolution preceding avian flight" "Science" 317:1378–1381 doi=10.1126/science.1144066
  53. ^
  54. ^ F. v. Huene. 1910. Über den ältesten Rest von Omosaurus (Dacentrurus) im englischen Dogger [On the oldest remains of Omosaurus (Dacentrurus) from the English Dogger]. Neues Jahrbuch für Mineralogie, Geologie und Paläontologie 1910(1):75-78.
  55. ^ O. Kuhn. 1964. Pars 105. Ornithischia (Supplementum I). In F. Westphal (ed.), Fossilium Catalogus. I: Animalia. IJssel Pers, Deventer, The Netherlands 1-80
  56. ^ Galton, P.M. and Powell, H.P., 1983, "Stegosaurian dinosaurs from the Bathonian (Middle Jurassic) of England, the earliest record of the family Stegosauridae", Geobios, 16: 219–229
  57. ^ Galton P.M. (1985) "British plated dinosaurs (Ornithischia, Stegosauridae), Journal of Vertebrate Paleontology, 5: 211-254
  58. ^ Galton, Peter M.; Upchurch, Paul (2004). "Stegosauria (Table 16.1)." In: Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press. Pp. 344-345. ISBN 0-520-24209-2.
  59. ^ P. M. Galton. 1990. Stegosauria. The Dinosauria, D. B. Weishampel, P. Dodson, & H. Osmólska (editors), University of California Press, Berkeley 435-455
  60. ^ S. C. R. Maidment, D. B. Norman, P. M. Barrett and P. Upchurch. 2008. Systematics and phylogeny of Stegosauria (Dinosauria: Ornithischia). Journal of Systematic Palaeontology 6(4):367-407
  61. ^ a b Ulansky, R. E., 2014. Evolution of the stegosaurs (Dinosauria; Ornithischia). Dinologia, 35 pp. [in Russian]. [DOWNLOAD PDF]
  62. ^ Dong Zhiming (1989). "On a small ornithopod (Gongbusaurus wucaiwanensis sp. nov.) from Kelamaili, Junggar Basin, Xinjiang, China". Vertebrata PalAsiatica. 27 (2): 140–146.
  63. ^ Norman, David B.; Sues, Hans-Dieter; Witmer, Larry M.; Coria, Rodolfo A. (2004). "Basal Ornithopoda". In Weishampel, David B.; Dodson, Peter; Osmólska Halszka (eds.). The Dinosauria (2nd ed.). Berkeley: University of California Press. pp. 393–412. ISBN 978-0-520-24209-8.
  64. ^ Knoll, Fabien (1999). "The family Fabrosauridae". In Canudo, J.I.; Cuenca-Bescós, G. (eds.). IV European Workshop on Vertebrate Palaeontology, Albarracin (Teruel, Spain), junio de 1999. Programme and Abstracts, Field guide. Servicio Publicaciones Universidad de Zaragoza. p. 54.
  65. ^ a b Fedak TJ. Description and evolutionary significance of the sauropodomorph dinosaurs from the Early Jurassic (Hettangian) McCoy Brook Formation. Ph.D. Dissertation, Dalhousie University. 2006.
  66. ^ a b Ulansky, R. E., 2014. Evolution of the stegosaurs (Dinosauria; Ornithischia). Dinologia, 35 pp. [in Russian]. [DOWNLOAD PDF]
  67. ^
  68. ^ Averianov, A.O.; Bakirov, A.A. & Martin, T. 2007. First definitive stegosaur from the Middle Jurassic of Kyrgyzstan. Paläontologische Zeitschrift 81(4):440–446.
  69. ^ Holtz, Thomas R. (1 Jan 2015). "Re: Phil Currie podcast interview + Jurassic dinosaurs". Dinosaur Mailing List.
  70. ^ Mortimer, Michael (2008). "Neotheropoda". The Theropod Database. Archived from the original on 2013-09-29. Retrieved 2009-11-17.
  71. ^ a b c Lambert, David; the Diagram Group (1990). The Dinosaur Data Book. New York: Avon Books. pp. 63–66, 250. ISBN 978-0-380-75896-8.
  72. ^ a b c d Dong, Zhiming; Y. Hasegawa; Y. Azuma (1990). The Age of Dinosaurs in Japan and China. Fukui, Japan: Fukui Prefectural Museum. p. 65 pp.
  73. ^ Sato, Tamaki; Hasegawa, Yoshikazu; Manabe, Makoto (2006). "A new elasmosaurid plesiosaur from the Upper Cretaceous of Fukushima, Japan". Palaeontology. 49 (3): 467–484. doi:10.1111/j.1475-4983.2006.00554.x.
  74. ^ Tsogtbaatar, K., D. Weishampel, D. C. Evans, and M. Watabe. (In review). A New Hadrosauroid (Dinosauria: Ornithopoda) from the Late Cretaceous Baynshire Formation of the Gobi Desert (Mongolia). PLOS ONE
  75. ^ Saito, Tsunemasa (1979). Wonder of the World's Dinosaurs. Tokyo: Kodansha Publishers. late 71.
  76. ^ Glut, Donald F. (1997). "Barsboldia". Dinosaurs: The Encyclopedia. Jefferson, North Carolina: McFarland & Co. p. 202. ISBN 978-0-89950-917-4.
  77. ^ Glut, Donald F. (1982). The New Dinosaur Dictionary. Secaucus, New Jersey: Citadel Press. p. 280. ISBN 978-0-8065-0782-8.
  78. ^ Lambert, David; the Diagram Group (1983). A Field Guide to Dinosaurs. New York: Avon Books. p. 153. ISBN 978-0-380-83519-5.
  79. ^ Norman, David B.; Sues, Hans-Dieter (2000). "Ornithopods from Kazakhstan, Mongolia and Siberia". In Benton, Michael J.; Shishkin, Mikhail A.; Unwin, David M.; Kurochkin, Evgenii N. (eds.). The Age of Dinosaurs in Russia and Mongolia. Cambridge: Cambridge University Press. pp. 462–479. ISBN 978-0-521-55476-3.
  80. ^ Kobayashi, Y.; Barsbold, R. (2006). "Ornithomimids from the Nemegt Formation of Mongolia" (PDF). Journal of the Paleontological Society of Korea. 22 (1): 195–207.
  81. ^ Malkani, M. Sadiq (20 May 2019). "Titanosaurian sauropod dinosaurs from Pakistan". ResearchGate. Retrieved 20 May 2019.
  82. ^ Malkani, M.S., 2015. Dinosaurs, Mesoeucrocodiles, Pterosaurs, New Fauna and Flora from Pakistan. Geological Survey of Pakistan, Information Release 823: 1-32.
  83. ^ Malkani, M.S., 2020. First Skull of Medium Sized Titanosaur in Indo-Pakistan Subcontinent Found from the Latest Maastrichtian Vitakri Formation of Pakistan; Associated Cranial and Postcranial Skeletons of Gspsaurus pakistani (Poripuchia, Stocky Titanosauria, Sauropoda) from Pakistan and India. Open Journal of Geology 10(4): 448-489.
  84. ^ a b Mortimer, M. (2018). "Ornithomimosauria". TheropodDatabase. Retrieved 2018-07-17.
  85. ^ Glut, Donald F. (2003). "Heilongjiangosaurus". Dinosaurs: The Encyclopedia. 3rd Supplement. Jefferson, North Carolina: McFarland & Company, Inc. p. 606. ISBN 978-0-7864-1166-5.
  86. ^ George Olshevsky. "Dinosaur Genera List update #180". Retrieved 2007-03-01.
  87. ^ Weirong, Li; Jidong, Jin (2001). "On the Upper Cretaceous Jiayin Group of Heilongjiang Province, China". In Deng Tao; Wang Yuan (eds.). Proceedings of the Eighth Annual Meeting of the Chinese Society of Vertebrate Paleontology (in Chinese and English). Beijing: China Ocean Press. pp. 65–74.
  88. ^ Hisa, K; Fukami, K; Murata, T; Shibuki, S; Haruyama, T; Tozawa, Y; Takeuchi, M; Sato, S; et al. (1988). "unknown" [A case of ileal hemorrhagic infarction of unknown origin (author's transl)]. Utan Scientific Magazine (in Japanese). 25 (8): 871–4. PMID 6968365. Cite uses generic title (help)
  89. ^ Zhiming, Dong; Y. Hasegawa; and Y. Azuma (1990). The Age of Dinosaurs in Japan and China. Fukui, Japan: Fukui Prefectural Museum.
  90. ^ M. Matsukawa and I. Obata. 1994. Dinosaurs and sedimentary environments in the Japanese Cretaceous: a contribution to dinosaur facies in Asia based on molluscan palaeontology and stratigraphy. Cretaceous Research 15(1):101-125
  91. ^ Novacek, Michael J. (1996). Dinosaurs of the Flaming Cliffs. New York: Anchor Books. ISBN 0-385-47774-0.
  92. ^ Norell, Mark A.; Makovicky, Peter J. (1999). "Important features of the dromaeosaurid skeleton II: information from newly collected specimens of Velociraptor mongoliensis". American Museum Novitates. 3282: 1–45. hdl:2246/3025.
  93. ^ a b c Olshevsky, G. (2019). "Dinosaur Genera List". Polychora. Retrieved 2019-11-02.
  94. ^
  95. ^ Robbins, Jim (21 January 2009). "Instead of Glory, the Finder of a Rare Dinosaur Fossil Faces Charges of Theft". The New York Times.
  96. ^ Missing or empty |title= (help)
  97. ^ "'Julieraptor' Dromaeosaurid Theropod Skeleton - Fossil Replica".
  98. ^ Mortimer, M. (2018). "Neotheropoda". TheropodDatabase. Retrieved 2018-07-18.
  99. ^ Dong, Z. (1973). "Dinosaurs from Wuerho". Reports of Paleontological Expedition to Sinkiang (II): Pterosaurian Fauna from Wuerho, Sinkiang. Memoirs of the Institute of Vertebrate Paleontology and Paleoanthropology, Academia Sinica 11: 45-52.
  100. ^ Grady, 1993. The Dinosaur Project: The Story of the Greatest Dinosaur Expedition Ever Mounted. Edmonton: Ex Terra Foundation; Toronto: Macfarlane Walter & Ross. ISBN 0-921912-46-3. 261 pp.
  101. ^
  102. ^ a b Harris, Jerald D. (2007-06-27). "Arsenic and Old Papers". Dinosaur Mailing List Archives. Cleveland Museum of Natural History. Retrieved 2007-07-29.
  103. ^ Kim, 1993. Journal of Natural History and Environments 1(1). World Society of Natural History and Environments:Pusan University, Pusan, Korea. ISSN 1225-6404.
  104. ^ Barrett, P M. 1999. A sauropod dinosaur from the Lower Lufeng Formation (Lower Jurassic) of Yunnan Province, People's Republic of China. Journal of Vertebrate Paleontology 19(4):785-787.
  105. ^ Dong, 1992. Dinosaurian Faunas of China. China Ocean Press (Beijing). 192 pp.
  106. ^ Mortimer, Mickey (2 January 2011). "The Theropod Database Blog: Sauropod Vertebrate Pictures Of the Year- Zhao's nomina nuda". Retrieved 1 January 2019.
  107. ^ a b Anonymous (2001). Chongqing Natural History Museum guidebook.
  108. ^ a b "Dinosaur Genera List corrections #171". Retrieved 1 January 2019.
  109. ^ Ellenberger, 1970. Les niveaux paléontologiques de première apparition des mammifères primoridaux en Afrique du Sud et leur ichnologie. Establissement de zones stratigraphiques detaillees dans le Stormberg du Lesotho (Afrique du Sud) (Trias Supérieur à Jurassique) [The paleontological levels of the first appearance of primordial mammals in southern Africa and their ichnology. Establishment of detailed stratigraphic zones in the Stormberg of Lesotho (southern Africa) (Upper Triassic to Jurassic). in Haughton (ed.). Second Symposium on Gondwana Stratigraphy and Paleontology, International Union of Geological Sciences. Council for Scientific and Industrial Research, Pretoria. 343-370.
  110. ^ Ellenberger and Ginsberg, 1966. Le gisement de Dinosauriens triasiques de Maphutseng (Basutoland) et l'origine des Sauropodes [The Triassic dinosaur locality of Maphutseng (Basutoland) and the origin of sauropods]. Comptes Rendus de l'Académie des Sciences à Paris, Série D. 262, 444-447.
  111. ^ Ellenberger, 1972. Contribution à la classification des Pistes de Vertébrés du Trias: Les types du Stormberg d'Afrique du Sud (I). Palaeovertebrata. 104, 152 pp.
  112. ^ Kitching and Raath, 1984. Fossils from the Elliot and Clarens Formations (Karoo Sequence) of the Northeastern Cape, Orange Free State and Lesotho, and a suggested biozonation based on tetrapods. Palaeontologia Africana. 25, 111-125.
  113. ^ Knoll, 2004. Review of the tetrapod fauna of the "Lower Stormberg Group" of the main Karoo Basin (southern Africa): Implication for the age of the Lower Elliot Formation. Bulletin de la Société Géologique de France. 175(1), 73-83.
  114. ^ Kranz, P. (1996). Notes on the sedimentary iron ores of Maryland and their dinosaurian faunas. Maryland Geological Survey Special Publications 3:87-115.
  115. ^ Chinnery, B.J., Lipka, T.R., Kirkland, J.I., Parrish, J.M., and Brett-Surman, M.K. (1998). Neoceratopsian teeth from the Lower Cretaceous of North America. In: Kirkland, J.I., and Estep, J.W. (eds.). Lower Cretaceous Terrestrial Ecosystems. New Mexico Museum of Natural History and Science Bulletin 14:297-302.
  116. ^ M. Sadiq Malkani;SUN Ge (2016) Fossil biotas from Pakistan with focus on dinosaur distributions and discussion on paleobiogeographic evolution of Indo-Pak Peninsula. Global Geology 19(4): 230-240
  117. ^ Zhao X. (1985). "Phylogeny and evolutionary stages of Dinosauria." Acta Palaeontologica Polonica 28(1-2); 295-306.
  118. ^ Mortimer, M (2004) "The Theropod Database" Archived 2013-09-29 at the Wayback Machine. University of Washington. Retrieved 2007-07-04.
  119. ^ Pickering, S., 1995. "Jurassic Park: Unauthorized Jewish Fractals in Philopatry", A Fractal Scaling in Dinosaurology Project, 2nd revised printing, Capitola, California: 478 pp.
  120. ^ "Re: What are these dinosaurs?". Retrieved 1 January 2019.
  121. ^ Lambert, D., and the Diagram Group. (1990). The Dinosaur Data Book . Facts on File: Oxford, England, 320 p.
  122. ^ Hasegawa Y, Manabe M, Hanai T, Kase T, Oji T. 1991. A diplodocoid dinosaur from the Early Cretaceous Miyako Group of Japan. Bulletin of the National Science Museum, Tokyo, Series C 17: 1–9.
  123. ^ Barrett PM, Hasegawa Y, Manabe M, Isaji S, Matsuoka H. 2002. Sauropod dinosaurs from the Lower Cretaceous of eastern Asia: taxonomic and biogeographical implications. Palaeontology 45: 1197–1217.
  124. ^ Azuma Y, Tomida Y. 1998. Japanese dinosaurs. In: Curie PJ, Padian K, eds. Encyclopaedia of dinosaurs. San Diego: Academic Press, 375–379.
  125. ^ Newton, E.T. (1899). On a megalosaurid jaw from Rhaetic beds near Bridgend (Glamorganshire). Quarterly Journal of the Geological Society of London 55:89-96.
  126. ^ "Dinosaur Genera List corrections #126". Retrieved 1 January 2019.
  127. ^ "Newtonsaurus et al". Retrieved 1 January 2019.
  128. ^ Zhao, X. (1983). "Phylogeny and evolutionary stages of Dinosauria." Acta Palaeontologica Polonica 28(1-2); 295-306.
  129. ^ Zhao, X. (1985). [The Jurassic Reptilia]. [The Jurassic System of China. Stratigraphy of China, No. 11] 286-290.
  130. ^ Weishampel, David B; et al. (2004). "Dinosaur distribution (Middle Jurassic, Asia)." In: Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd Edition, Berkeley: University of California Press. Pp. 541–542. ISBN 0-520-24209-2.
  131. ^ Malkani, M. "Titanosaurian sauropod dinosaurs from Pakistan". ResearchGate. Retrieved 2019-05-23.
  132. ^ Dong, Z. (1992). Dinosaurian Faunas of China. China Ocean Press. pp. 111–113. ISBN 3-540-52084-8.
  133. ^ Rothschild, B.M.; Molnar, R.E. (2005). "Sauropod Stress Fractures as Clues to Activity". In Tidwell, Virginia; Carpenter, Kenneth (eds.). Thunder-Lizards, The Sauropodomorph Dinosaurs. Indiana University Press. pp. 381–392. ISBN 0-253-34542-1.
  134. ^ Triebold, M. (1997). "The Sandy site: Small dinosaurs from the Hell Creek Formation of South Dakota." in Wolberg, D., Stump, E. and Rosenberg, G. (eds); Dinofest International: Proceedings of a Symposium. Arizona State University Academy of Natural Science. 245-48
  135. ^ Mortimer, Mickey (2011-01-02). "Sauropod Vertebrate Pictures Of the Year- Zhao's nomina nuda". Retrieved 2 January 2011.
  136. ^ Dingus, L.; Norrell, M.A. (2010). "The Canadian dinosaur bone rush". Barnum Brown: The man who discovered Tyrannosaurus rex. University of California Press. p. 137. ISBN 978-0-520-25264-6.
  137. ^ a b Gee, H. & Rey, L. V. 2003. A Field Guide to Dinosaurs: The Essential Handbook for Travelers in the Mesozoic. Barron's Educational Series, Inc.
  138. ^ page 9 of
  139. ^ a b Delair, J.B., and Sarjeant, W.A.S. (2002). The earliest discoveries of dinosaurs: the records re-examined. Proceedings of the Geologists' Association 113:185–197.
  140. ^ Lhuyd, E. (1699). Lithophylacii Britannici Ichnographia, sive lapidium aliorumque fossilium Britannicorum singulari figura insignium. Gleditsch and Weidmann:London.
  141. ^ Gunther, R.T. (1945). Early Science in Oxford: Life and Letters of Edward Lhuyd, volume 14. Author:Oxford.
  142. ^ a b Kirkland, J.I.; Hernández-Rivera, R.; Gates, T.; Paul, G.S.; Nesbitt, S.; Serrano-Brañas, C.I.; Garcia-de la Garza, J.P. (2006). "Large hadrosaurine dinosaurs from the latest Campanian of Coahuila, Mexico". In Lucas, S.G.; Sullivan, R.M. (eds.). Late Cretaceous Vertebrates from the Western Interior. New Mexico Museum of Natural History and Science Bulletin. 35. Albuquerque: New Mexico Museum of Natural History and Science. pp. 299–315.
  143. ^ Prieto-Márquez, A. (2014). "Skeletal morphology of Kritosaurus navajovius (Dinosauria: Hadrosauridae) from the Late Cretaceous of the North American south-west, with an evaluation of the phylogenetic systematics and biogeography of Kritosaurini". Journal of Systematic Palaeontology. 12 (2): 133–175. doi:10.1080/14772019.2013.770417. S2CID 84942579.
  144. ^ a b Peter M. Galton; Kenneth Carpenter (2016). "The plated dinosaur Stegosaurus longispinus Gilmore, 1914 (Dinosauria: Ornithischia; Upper Jurassic, western USA), type species of Alcovasaurus n. gen". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 279 (2): 185–208. doi:10.1127/njgpa/2016/0551.
  145. ^ Ulansky, R. E., 2014. Evolution of the stegosaurs (Dinosauria; Ornithischia). Dinologia, 35 pp. [in Russian].
  146. ^ "Sanchusaurus (Dino Kingdom)". Nakasato Dinosaur Kingdom. kanna Dinosaur Center. Archived from the original on 2007-10-05. Retrieved 2008-01-10.
  147. ^ Worth, Graeme. "SANCHUSAURUS". The Dinosaur Encyclopaedia 4.0. HyperWorks Reference Software. Archived from the original on October 4, 2007. Retrieved 2012-12-07.
  148. ^ Malkani, S.M. (2015). "Titanosaurian sauropod dinosaurs from Pakistan". Proceedings of the 12th Symposium of "Mesozoic Terrestrial Ecosystems (MTE 12). Abstract volume: 93–98.
  149. ^ Tykoski, R. S. 1997. A new ceratosaurid theropod from the Early Jurassic Kayenta Formation of northern Arizona. Journal of Vertebrate Paleontology 17 (3, Supplement):81A.
  150. ^ Tykoski, R. S. 1998. The osteology of Syntarsus kayentakatae and its implications for ceratosaurid phylogeny. Master's thesis, The University of Texas at Austin, Austin, Texas, 217 pp.
  151. ^ Ulansky, R. E., 2014. Evolution of the stegosaurs (Dinosauria; Ornithischia). Dinologia, 35 pp. [in Russian]. PDF.
  152. ^ Lyon, 2000.
  153. ^ Sereno, Wilson and Conrad, 2004. New dinosaurs link southern landmasses in the Mid-Cretaceous. Proceedings of the Royal Society, Series B. 271, 1325-1330.
  154. ^ Sereno and Brusatte, 2008. Basal abelisaurid and carcharodontosaurid theropods from the Lower Cretaceous Elrhaz Formation of Niger. Acta Palaeontologica Polonica. 53(1), 15-46.
  155. ^ Sereno, 2010. Noasaurid (Theropoda: Abelisauroidea) skeleton from Africa shows derived skeletal proportions and function. Journal of Vertebrate Paleontology. Program and Abstracts 2010, 162A.
  156. ^ Ulansky, R. E., 2014. Evolution of the stegosaurs (Dinosauria; Ornithischia). Dinologia, 35 pp. [in Russian]. [DOWNLOAD PDF]
  157. ^ Boyd, Clint Aaroen (May 2012). "Taxonomic revision of latest Cretaceous North american basal neonithischian taxa and a phylogenetic analysis of basal ornithischian relationships". Cite journal requires |journal= (help)
  158. ^ "Cretaceous Crocs Crunched Little Dinosaurs". Science. 27 February 2013. Retrieved 30 September 2020.
  159. ^ a b Chris Hansen (March 30, 2019), Lawmakers could hear proposal to name state's official dinosaur, iFiberOne News
  160. ^ a b Legislature weighs proposal for official state dinosaur, (KIRO (AM)), March 25, 2019
  161. ^ Peecook, Brandon R.; Sidor, Christian A.; Evans, David C. (20 May 2015). "The First Dinosaur from Washington State and a Review of Pacific Coast Dinosaurs from North America". PLOS ONE. 10 (5): e0127792. Bibcode:2015PLoSO..1027792P. doi:10.1371/journal.pone.0127792. PMC 4439161. PMID 25993090.
  162. ^ "Burke Museum paleontologists discover the first dinosaur fossil in Washington state". Seattle: Burke Museum . May 19, 2015.
  163. ^ Parkland 4th graders petitioned for 'Suciasaurus rex' to be Washington state dinosaur – If Washington gets a state dinosaur, residents can thank students from Parkland's Elmhurst Elementary., KING-TV, April 7, 2019
  164. ^ Dong, Hasegawa and Azuma, 1990. The Age of Dinosaurs in Japan and China. Fukui, Japan: Fukui Prefectural Museum. 65 pp.
  165. ^ Azuma, 1991. Early Cretaceous Dinosaur Fauna from the Tetori Group, central Japan. Research on Dinosaurs from the Tetori Group (1). Professor S. Miura Memorial Volume, 55-69.
  166. ^ "Macronaria". Retrieved 1 January 2019.
  167. ^ Yun, Chan-gyu (2017). "Teihivenator gen. nov., a new generic name for the Tyrannosauroid Dinosaur "Laelaps" macropus (Cope, 1868; preoccupied by Koch, 1836)". Journal of Zoological and Bioscience Research. 4.
  168. ^ Marjanovic, D. (2020). "Re: [dinosaur] "Yunyangosaurus" is not available". Dinosaur Mailing List. Clevelant Museum of Natural History. Retrieved 2020-01-25.
  169. ^ Brownstein, C.D. "Theropod specimens from the Navesink Formation and their implications for the Diversity and Biogeography of Ornithomimosaurs and Tyrannosauroids on Appalachia". PeerJ Preprints. 5: e3105v1. doi:10.7287/peerj.preprints.3105v1.
  170. ^ Brownstein, C.D. (2018). "A tyrannosauroid tibia from the Navesink Formation of New Jersey and its biogeographic and evolutionary implications for North American tyrannosauroids". Cretaceous Research. 305: 309–318. doi:10.1016/j.cretres.2018.01.005.
  171. ^ "Therizinosauroidea". Retrieved 1 January 2019.
  172. ^ "Re: Tiantai(o)saurus Dong et al. 2007, missed Chinese therizinosaur?". Retrieved 1 January 2019.
  173. ^ "Re: Tonouchisaurus". Retrieved 1 January 2019.
  174. ^ Blows, W.T., 1998, "A review of Lower and Middle Cretaceous dinosaurs of England", New Mexico Museum of Natural History and Science Bulletin, Albuquerque 14: 29-38
  175. ^ Martill, D.M. and Naish, D., 2001, "Ornithopod dinosaurs." Pages 60-132 in Martill, D.M. and Naish, D. (eds.). Dinosaurs of the Isle of Wight, The Palaeontological Association, London.
  176. ^ Galton, P.M., 2009, "Notes on Neocomian (Late Cretaceous) ornithopod dinosaurs from England - Hypsilophodon, Valdosaurus, "Camptosaurus", "Iguanodon" - and referred specimens from Romania and elsewhere", Revue de Paléobiologie 28(1): 211-273
  177. ^ Delair, J.B., 1982, "Notes on an armoured dinosaur from Barnes High, Isle of Wight", Proceedings of the Isle of Wight Natural History and Archaeological Society for 1980, 7(5): 297-302
  178. ^ Blows W.T. (1987). The armoured dinosaur Polacanthus foxi, from the Lower Cretaceous of the Isle of Wight, Palaeontology. 30, 557–580
  179. ^ Malkani, M.S. (2006). "First rostrum of carnivorous Vitakridrinda (Abelisaurid theropod dinosaur) found from the Latest Cretaceous dinosaur beds (Vitakri) Member of Pab Formation, Alam Kali Kakor Locality of Viakri area, Darkham District, Balochistan, Pakistan". Sindh University y Research Journal (Science Series). 38 (3): 7–26.
  180. ^ Malkani M.S. 2015a. Terrestrial mesoeucrocodiles from the Cretaceous of Pakistan. In: Zhang Y., Wu S.Z., Sun G. eds., abstract volume, 12th Symposium on “Mesozoic Terrestrial Ecosystems (MTE 12), and 3rd Symposium of International Geoscience Program (IGCP 608) “Cretaceous Ecosystem of Asia and Pacific” August 15–20, 2015, Paleontological Museum of Liaoning/Shenyang Normal University, Shenyang, China, 242-246.
  181. ^ Malkani, M.S., 2015b. Dinosaurs, mesoeucrocodiles, pterosaurs, new fauna and flora from Pakistan. Geological Survey of Pakistan, Information Release 823: i-iii,1-32.
  182. ^ Holtz, Thomas R. (2012). "Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages" (PDF).
  183. ^ a b
  184. ^ Olshevsky, George (December 4, 2000). "Dinosaur Genera List". Retrieved July 18, 2018.
  185. ^ Mortimer, Mickey (March 26, 2011). "The Theropod Database". Sauropodomorph cladogram. Retrieved July 18, 2018.
  186. ^ Missing or empty |title= (help)
  187. ^ Li, Zhengqi. (2001). Distribution, burying and classification of dinosaur fossils in Upper Cretaceous strata at Meipu Town, Yunxian County of Hubei Province. Hubei Geology & Mineral Resources, 15(4)(Total No 37): 25-31. [1]
  188. ^ Zhou, S.Q.D. (2005). The Dinosaur Egg Fossils in Nanyang, China. China University of Geosciences Press. pp. 1–145. ISBN 978-7-562-52033-7.
  189. ^ Barsbold, R., Maryanska, T., and Osmolska, H. (1990). "Oviraptorosauria," in Weishampel, D.B., Dodson, P., and Osmolska, H. (eds.). The Dinosauria. Berkeley: University of California Press, pp. 249-258.
  190. ^ Clark, J.M., Norell, M.A., & Barsbold, R. (2001). "Two new oviraptorids (Theropoda:Oviraptorosauria), upper Cretaceous Djadokhta Formation, Ukhaa Tolgod, Mongolia." Journal of Vertebrate Paleontology 21(2):209-213., June 2001.
  191. ^ K. Li, Y. Zhang, K. Cai, 1999, The Characteristics of the Composition of the Trace Elements in Jurassic Dinosaur Bones and Red Beds in Sichuan Basin, Geological Publishing House, Beijing

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